Emergent features, attention, and object perception

The perceptual processing of arrows and triangles and of their component angles and lines was explored in a number of different tasks. The results suggest that some analysis of shapes into simpler parts occurs preattentively, because these parts can recombine to form illusory conjunctions when attention is divided. The presence of "emergent features," such as closure or arrow junctions, was inferred from predicted correlations in the pattern of performance across tasks and across individual subjects. Thus triangles (for most subjects) and arrows (for some subjects) behave as if they had a perceptual feature that is absent from their parts and that mediates parallel detection in search and easy texture segregation. For some subjects, circles could apparently supply the additional feature (presumably closure) required to form illusory triangles from their component lines, whereas for other subjects circles had no effect. The fact that triangle lines can form illusory conjunctions with another shape makes it unlikely that triangles are perceived holistically and strengthens the interpretation that relies on emergent features.     

Perceptual grouping and attention in visual search for features and for objects

This article explores the effects of perceptual grouping on search for targets defined by separate features or by conjunction of features. Treisman and Gelade proposed a feature-integration theory of attention, which claims that in the absence of prior knowledge, the separable features of objects are correctly combined only when focused attention is directed to each item in turn. If items are preattentively grouped, however, attention may be directed to groups rather than to single items whenever no recombination of features within a group could generate an illusory target. This prediction is confirmed: In search for conjunctions, subjects appear to scan serially between groups rather than items. The scanning rate shows little effect of the spatial density of distractors, suggesting that it reflects serial fixations of attention rather than eye movements. Search for features, on the other hand, appears to independent of perceptual grouping, suggesting that features are detected preattentively. A conjunction target can be camouflaged at the preattentive level by placing it at the boundary between two adjacent groups, each of which shares one of its features. This suggests that preattentive grouping creates separate feature maps within each separable dimension rather than one global configuration.     

Relationship between visual binding, reentry and awareness

Visual feature binding has been suggested to depend on reentrant processing. We addressed the relationship between binding, reentry, and visual awareness by asking the participants to discriminate the color and orientation of a colored bar (presented either alone or simultaneously with a white distractor bar) and to report their phenomenal awareness of the target features. The success of reentry was manipulated with object substitution masking and backward masking. The results showed that late reentrant processes are necessary for successful binding but not for phenomenal awareness of the bound features. Binding errors were accompanied by phenomenal awareness of the misbound feature conjunctions, demonstrating that they were experienced as real properties of the stimuli (i.e., illusory conjunctions). Our results suggest that early preattentive binding and local recurrent processing enable features to reach phenomenal awareness, while later attention-related reentrant iterations modulate the way in which the features are bound and experienced in awareness.     

The role of attention in illusory conjunctions

In five experiments, we investigated the effects of attention on illusory conjunctions formed between features of unrelated objects. The first three experiments used a weak manipulation of attention and found that illusory conjunctions formed either among features receiving high attentional priority or among features receiving low attentional priority were not more frequent than were conjunctions formed between mixed features of different attentional priority. The last two experiments used a strong manipulation of attention and failed to reveal any evidence of true illusory conjunctions. The results are inconsistent with the feature-integration theory, which predicts that when attention is focused on a subset of items, illusory conjunctions ought to occur within and outside of the attended subset, but not between the attended and unattended items.     

When do letter features migrate? A boundary condition for feature-integration theory

Feature-integration theory postulates that a lapse of attention will allow letter features to change position and to recombine as illusory conjunctions (Treisman & Paterson, 1984). To study such errors, we used a set of uppercase letters known to yield illusory conjunctions in each of three tasks. The first, a bar-probe task, showed whole-character mislocations but not errors based on feature migration and recombination. The second, a two-alternative forced-choice detection task, allowed subjects to focus on the presence or absence of subletter features and showed illusory conjunctions based on feature migration and recombination. The third was also a two-alternative forced-choice detection task, but we manipulated the subjects' knowledge of the shape of the stimuli: In the case-certain condition, the stimuli were always in uppercase, but in the case-uncertain condition, the stimuli could appear in either upper- or lowercase. Subjects in the case-certain condition produced illusory conjunctions based on feature recombination, whereas subjects in the case-uncertain condition did not. The results suggest that when subjects can view the stimuli as feature groups, letter features regroup as illusory conjunctions; when subjects encode the stimuli as letters, whole items may be mislocated, but subletter features are not. Thus, illusory conjunctions reflect the subject's processing strategy, rather than the architecture of the visual system.     

Functional theory of illusory conjunctions and neon colors

Illusory conjunctions are the incorrect perceptual combination of briefly presented colors and shapes. In the neon colors illusion, achromatic figures take on the color of an overlaid grid of colored lines. Both illusions are explained by a theory that assumes (a) poor location information or poor spatial resolution for some aspects of visual information and (b) that the spatial location of features is constrained by perceptual organization. Computer simulations demonstrate that the mechanisms suggested by the theory are useful in veridical perception and they are sufficient to produce illusory conjunctions. The theory suggests mechanisms that economically encode visual information in a way that filters noise and fills in missing data. Issues related to neural implementation are discussed. Four experiments illustrate the theory. Illusory conjunctions are shown to be affected by objective stimulus organization, by subjective organization, and by the linguistic structure of ambiguous Hebrew words. Neon colors are constrained by linguistic structure in the same way as illusory conjunctions.     

Conjunction of color and form without attention: evidence from an orientation-contingent color aftereffect

According to feature-integration theory (Treisman & Gelade, 1980), separable features such as color and shape exist in separate maps in preattentive vision and can be integrated only through the use of spatial attention. Many perceptual aftereffects, however, which are also assumed to reflect the features available in preattentive vision, are sensitive to conjunctions of features. One possible resolution of these views holds that adaptation to conjunctions depends on spatial attention. We tested this proposition by presenting observers with gratings varying in color and orientation. The resulting McCollough aftereffects were independent of whether the adaptation stimuli were presented inside or outside of the focus of spatial attention. Therefore, color and shape appear to be conjoined preattentively, when perceptual aftereffects are used as the measure. These same stimuli, however, appeared to be separable in two additional experiments that required observers to search for gratings of a specified color and orientation. These results show that different experimental procedures may be tapping into different stages of preattentive vision.     

Illusory conjunctions and the cerebral hemispheres

Two visual half-field experiments tested Moscovitch’s (1979) proposition that cerebral asymmetry does not concern the earliest perceptual stages but only later processing. Subjects were briefly shown displays that included one (Experiment 1) or two (Experiment 2) types of forms differing in size and which, according to previous evidence, might lead to opposite laterality effects. Laterality effects were assessed for correct detections and for illusory conjunctions, both in terms of raw detection scores and in terms of perceptual discriminability (dr scores). In Experiment 1, displays included either rectangles or triangles. In the first case, the target was a cross; in the second case, it was a Star of David. A hemifield x size interaction was observed both on correct detections and on associated discriminability. Yet, no such interaction was obtained for illusory conjunctions or for associated d’ scores. In Experiment 2, the two types of forms were presented simultaneously, with the small ones either inside or outside the large ones. No laterality effects were observed. Some implications of these data for both hemispheric asymmetry and feature integration issues are discussed. The results suggest that early preattentive processes of feature extraction are not lateralized, whereas some integrative mechanisms, such as Treisman’s (1988) focal attention, may operate differently in the two hemispheres.     

Integration of local features as a function of global goodness and spacing

In two experiments, the accuracy with which subjects detected a conjunction of features was examined as a function of the spacing between items and the goodness of the axis along which they were located. In each array, two items were arranged along a vertical, a horizontal, or a diagonal axis. Based on the well-established oblique effect, the vertical and horizontal axes were considered to be good global patterns and the diagonals were considered to be poor. In Experiment 1, the two items in an array could be two horizontal lines, two vertical lines, a vertical and a horizontal line, or a plus sign with one of the single lines. In Experiment 2, a positive- and a negative-diagonal line were used as the individual features, and an "X" was used as the conjunction. The results from Experiment 1 indicated that global goodness influenced only the rate of illusory conjunctions, and not of feature errors. Illusory conjunctions of vertical and horizontal line segments were more likely to occur in vertical and horizontal arrangements. The results from Experiment 2 revealed a reversal of the effect of global goodness on the rate of illusory conjunctions: Illusory conjunctions of negative- and positive-diagonal line segments were more likely to occur in diagonal arrangements. The results of both experiments taken together showed the existence of an important and new factor that influences the likelihood that features of shape will be conjoined: the ease with which line segments conjoin when they are translated along their extent toward each other. In both experiments, greater spacing between items produced more feature-identification errors and fewer feature-integration errors than did less spacing.     

On the Reality of Illusory Conjunctions

The reality of illusory conjunctions in perception has been sometimes questioned, arguing that they can be explained by other mechanisms. Most relevant experiments are based on migrations along the space dimension. But the low rate of illusory conjunctions along space can easily hide them among other types of errors. As migrations over time are a more frequent phenomenon, illusory conjunctions can be disentangled from other errors. We report an experiment in which series of colored letters were presented in several spatial locations, allowing for migrations over both space and time. The distribution of frequencies were fit by several multinomial tree models based on alternative hypothesis about illusory conjunctions and the potential sources of free-floating features. The best-fit model acknowledges that most illusory conjunctions are migrations in the time domain. Migrations in space are probably present, but the rate is very low. Other conjunction errors, as those produced by guessing or miscategorizations of the to-be-reported feature, are also present in the experiment. The main conclusion is that illusory conjunctions do exist.     

Effect of feature similarity on illusory conjunctions

In four experiments, we examined whether the phenomenon of illusory conjunctions is constrained by feature similarity. Specifically, are illusory conjunctions more likely to occur between items with similar features than between items with dissimilar features? Feature similarity was manipulated in two dimensions: color and shape. Experiment 1 demonstrated that more illusory conjunctions occur between items with similar colors than between items with dissimilar colors. A similar effect was found for letter similarity in Experiment 2. Experiments 3 and 4 demonstrated that the similarity effect is still obtained for illusory conjunctions even when identification of the relevant features is near perfect. These findings introduce a new constraint in the process of feature integration. Additional implications of the similarity constraint for theories of feature integration are discussed.     

Illusory conjunctions in the perception of objects

In perceiving objects we may synthesize conjunctions of separable features by directing attention serially to each item in turn (A. Treisman and G. Gelade, Cognitive Psychology, 1980, 12, 97–136). This feature-integration theory predicts that when attention is diverted or overloaded, features may be wrongly recombined, giving rise to “illusory conjunctions.” The present paper confirms that illusory conjunctions are frequently experienced among unattended stimuli varying in color and shape, and that they occur also with size and solidity (outlined versus filled-in shapes). They are shown both in verbal recall and in simultaneous and successive matching tasks, making it unlikely that they depend on verbal labeling or on memory failure. They occur as often between stimuli differing on many features as between more similar stimuli, and spatial separation has little effect on their frequency. Each feature seems to be coded as an independent entity and to migrate, when attention is diverted, with few constraints from the other features of its source or destination.     

Illusory conjunctions are alive and well: a reply to Donk (1999).

When presented with a red T and a green O, observers occasionally make conjunction responses and indicate that they saw a green T. These errors have been interpreted as reflecting separable processing stages of feature detection and integration with the illusory conjunctions arising from a failure at the integration stage. Recently, M. Donk (1999) asserted that the phenomenon of illusory conjunctions is an artifact. Conjunction reports are actually the result of confusing a nontarget item (O in the example above) for a target item (the letter T) and (correctly) reporting the color associated with the (incorrectly) selected target. The authors demonstrate that although target-nontarget confusion errors are a potential source of conjunction reports, there is a plethora of findings that cannot be accounted for by this confusion model. A review of the literature indicates that in many studies, illusory conjunctions do result from a failure to properly integrate features.     

Illusory conjunctions reflect the time course of the attentional blink

Illusory conjunctions in the time domain are binding errors for features from stimuli presented sequentially but in the same spatial position. A similar experimental paradigm is employed for the attentional blink (AB), an impairment of performance for the second of two targets when it is presented 200–500 msec after the first target. The analysis of errors along the time course of the AB allows the testing of models of illusory conjunctions. In an experiment, observers identified one (control condition) or two (experimental condition) letters in a specified color, so that illusory conjunctions in each response could be linked to specific positions in the series. Two items in the target colors (red and white, embedded in distractors of different colors) were employed in four conditions defined according to whether both targets were in the same or different colors. Besides the U-shaped function for hits, the errors were analyzed by calculating several response parameters reflecting characteristics such as the average position of the responses or the attentional suppression during the blink. The several error parameters cluster in two time courses, as would be expected from prevailing models of the AB. Furthermore, the results match the predictions from Botella, Barriopedro, and Suero’s (Journal of Experimental Psychology: Human Perception and Performance, 27, 1452–1467, 2001) model for illusory conjunctions.     

视觉特征捆绑的认知机制研究

以大学生为被试,探讨视觉特征捆绑的认知机制。实验采用空间提示范式,要求被试完成字母形状和颜色的辨别任务,考察集中注意和分散注意条件下的IC差异。结果表明,注意在视觉特征捆绑中起重要作用,集中注意能有效地减少IC的出现。同时注意对特征捆绑的调节作用表现出距离效应,即近距离比远距离的IC错误多,提示了注意的作用机制有可能与空间距离信息有关。     

Illusory conjunctions inside and outside the focus of attention

This article addresses 2 questions that arise from the finding that visual scenes are first parsed into visual features: (a) the accumulation of location information about objects during their recognition and (b) the mechanism for the binding of the visual features. The first 2 experiments demonstrated that when 2 colored letters were presented outside the initial focus of attention, illusory conjunctions between the color of one letter and the shape of the other were formed only if the letters were less than 1 degree apart. Separation greater than 2 degrees resulted in fewer conjunction errors than expected by chance. Experiments 3 and 4 showed that inside the spread of attention, illusory conjunctions between the 2 letters can occur regardless of the distance between them. In addition, these experiments demonstrated that the span of attention can expand or shrink like a spotlight. The results suggest that features inside the focus of attention are integrated by an expandable focal attention mechanism that conjoins all features that appear inside its focus. Visual features outside the focus of attention may be registered with coarse location information prior to their integration. Alternatively, a quick and imprecise shift of attention to the periphery may lead to illusory conjunctions among adjacent stimuli.     

Is meaning implicated in illusory conjunctions?

According to feature-integration theory, when attention is diverted from a display, features from different objects in that display may be wrongly recombined, giving rise to "illusory conjunctions" (Treisman & Schmidt, 1982). Two experiments are reported that examine the nature of these illusory conjunctions. In displays that contain color names and adjectives printed in colored ink, subjects made two kinds of interesting and previously unreported errors. Consider, for example, a display that included the word BROWN in red ink and the word HEAVY in green ink. Subjects would sometimes incorrectly report that the word RED or the ink color brown had appeared in the display (e.g., RED in green ink or HEAVY in brown ink). It appears that subjects extract semantic representations from input and are sometimes confused about whether a particular representation has been extracted from a word or a color patch. Contrary to feature-integration theory, these findings suggest that illusory conjunctions may occur with high-level codes as well as with perceptual features.     

Illusory conjunctions die hard: a reply to Prinzmetal, Diedrichsen, and Ivry (2001).

M. Donk (1999) showed that various data patterns that have been considered as evidence for the existence of illusory conjunctions may be due to errors of target-nontarget confusion, an account that challenges the mere existence of illusory conjunction. In a reply, W. Prinzmetal, J. Diedrichsen, and R. B. Ivry (2001) argued against this conclusion, claiming that some earlier findings can be explained only when one assumes that illusory conjunctions exist. The current article shows that Prinzmetal et al.'s claims cannot refute any of Donk's earlier conclusions, suggesting indeed that one can only conclude that "illusory conjunctions are an illusion."     

Illusory Conjunctions of Color and Motion With Shape Following Bilateral Parietal Lesions

The primate visual system responds to shapes, colors, and various other features more strongly in some brain areas than others. However, it remains unclear how these features are bound together so that an object with all its attributes is perceived. A patient (R.M.) with bilateral parietal-occipital lesions has been shown previously to miscombine the shape and color of items, making errors known as illusory conjunctions (ICs). In this study, we examined the effects of a third feature (motion) on this patient's IC rates. R.M. was presented with two letters that moved in different ways. He often reported seeing the shape of one of the letters with the other letter's motion. His performance on the same task with three features shows that correctly combining two features did not necessarily lead to correctly binding the third. These data support modularity of feature representations in the human brain and provide supporting evidence that spatial representations associated with the parietal lobe are necessary for normal feature integration.