Reflex augmentation of a tap-elicited eyeblink: the effects of tone frequency and tap intensity

The amplitude of the human eyeblink elicited by a mild tap between the eyebrows can be increased if a brief tone is presented simultaneously with the tap. In two experiments we examined how this reflex augmentation effect varies with changes in the frequency characteristics of the tone and with changes in the intensity of the tap. We also examined how these effects change in the course of development. For newborn infants and for adults, fixed intensity tones with frequencies of either 1, 2, or 4 kHz produced equivalent amounts of reflex augmentation. Furthermore, while blink amplitude was an increasing function of tap intensity in both populations, the amount of reflex augmentation engendered by a simultaneously presented tone was independent of the intensity of the tap.     

A comparison of chinchilla auditory evoked response and behavioral response thresholds

The auditory sensitivity of the chinchilla was measured by both a behavioral and auditory evoked-response (AER) procedure. There was no significant difference in either the waveform or the threshold of the AER recorded over the tentorium from awake and anesthetized chinchillas. The chinchilla threshold for pure tones appears to be more sensitive when measured by the behavioral technique than by the AER method. Both the AER and behavioral response of three chinchillas exposed to a 105 dB SPL octave band of noise centered at 2 KHz for 3 h shifted markedly after the exposure. However, the difference between the AER and behavioral response threshold was reduced or absent. During the process of recovery the difference between the two responses begins to reappear.     

A clarification of continuous repertoire development

The key-peck response of five pigeons was reinforced on a schedule whenever the interval between pecks at two response keys was between 1.0 and 2.33 seconds in the presence of a 2,500-Hertz tone or between 4.66 and 6.0 seconds in the presence of a 1250-Hertz tone. There was no tendency for responses of intermediate duration to occur when test tones of intermediate frequency were presented. This result clarifies a previous finding using a similar procedure but with a visual intensity stimulus dimension.     

Preference for tones as a function of frequency (hertz) and intensity (decibels)

An extension of the Wundt hypothesis concerning stimulus intensity and hedonic tone leads to the prediction that Ss prefer tones of intermediate frequency (hertz) and of intermediate intensity (decibels). In Experiment 1, tones varying from 60 to 5000 Hz were presented in a paired comparison procedure, and, as predicted, Ss′ preference was an inverted-U-shaped function of frequency, with the most preferred tones in the range of 400–750 Hz. In Experiment 2, tones varying in intensity from 40 to 90 dB were also presented in a paired comparison procedure, and again the predicted inverted-U-shaped preference function was found, with 50 dB as the preferred intensity. Related research and some unexpected findings are discussed.     

The developmental course of lexical tone perception in the first year of life

Perceptual reorganisation of infants' speech perception has been found from 6 months for consonants and earlier for vowels. Recently, similar reorganisation has been found for lexical tone between 6 and 9 months of age. Given that there is a close relationship between vowels and tones, this study investigates whether the perceptual reorganisation for tone begins earlier than 6 months. Non-tone language English and French infants were tested with the Thai low vs. rising lexical tone contrast, using the stimulus alternating preference procedure. Four- and 6-month-old infants discriminated the lexical tones, and there was no decline in discrimination performance across these ages. However, 9-month-olds failed to discriminate the lexical tones. This particular pattern of decline in nonnative tone discrimination over age indicates that perceptual reorganisation for tone does not parallel the developmentally prior decline observed in vowel perception. The findings converge with previous developmental cross-language findings on tone perception in English-language infants [Mattock, K., & Burnham, D. (2006). Chinese and English infants' tone perception: Evidence for perceptual reorganization. Infancy, 10(3)], and extend them by showing similar perceptual reorganisation for non-tone language infants learning rhythmically different non-tone languages (English and French).     

Behavior-Based Assessment of the Auditory Abilities of Brushtail Possums

Brushtail possums (Trichosurus vulpecula) were trained to press a right lever when a tone was presented (a tone‐on trial) and a left lever when a tone was not presented (a tone‐off trial) to gain access to food. During training the tone was set at 80 dB(A), with a frequency of 0.88 kH for 3 possums and of 4 kH for the other 2. Once accuracy was over 90% correct across five consecutive sessions, a test session was conducted where the intensity of the tone was reduced by 8 dB(A) over blocks of 20 trials until accuracy over a block fell below 60%. After each test session, training sessions were reintroduced and continued until accuracy was again over 90%, when another test session was conducted. This process continued until there were at least five test sessions at that tone frequency. The same procedure was then used with frequencies of 0.20, 0.88, 2, 4, 10, 12.5, 15, 20, 30, and 35 kHz. Percentage correct and d' decreased approximately linearly for all possums as tone intensity reduced. Both sets of lines were shallowest at the higher frequencies and steepest at the lower frequencies. Hit and false alarm rates mirrored each other at high frequencies but were asymmetric at lower frequencies. Equal d' contours showed that sensitivity increased from 2 to 15 kHz and continued to be high over 20 to 35 kHz. The possums remained sensitive to the 20 to 35 kHz tones even at low intensities. The present study is the first to report the abilities of possum to detect tones over this range of frequencies and its results support the findings of a microelectrode mapping survey of possums' auditory cortex.     

Perceptual grouping in audition

The present experiments evaluated the effect of relative frequency as a determinant of the figure-ground organization of sequences of auditory tones. Observers counted sequences of 20 ms tones that were presented at the same frequency or that alternated between two different frequencies. The alternating tones differed in frequency by one whole tone, seven tones, or nineteen tones. Counting accuracy increased with increases in the silent interval between the tones. When the alternating tones differed by seven or nineteen tones, counting was disrupted at rates of presentation of eight tones per second or slower. In contrast to this decrement in the counting of tones that alternated by over an octave, very little decrement was observed when the tones alternated by just one whole tone. The best subjects counted these alternating tones more accurately than the tones presented at the same frequency. The poorest subjects showed a small decrement even when the tones alternated by just one whole tone. The results were discussed in terms of determinants of figure-ground organization in auditory information processing.     

Does stimulus context affect loudness or only loudness judgments?

Marks (1988) reported that when equal-loudness matches were inferred from magnitude estimates of loudness for tones of two different frequencies, the matches were affected by changes in the stimulus intensity range at both frequencies. Marks interpreted these results as reflecting the operation of response biases in the subjects' estimates; that is, the effect of range was to alter subjects' judgments but not necessarily the perception of loudness itself. We investigated this effect by having subjects choose which of two tone pairs defined the larger loudness interval. By using tones of two frequencies, and varying their respective intensity ranges, we reproduced Marks' result in a procedure devoid of numerical responses. When the tones at one frequency are all soft, but the tones at the other frequency are not all soft, cross-frequency loudness matches are different from those obtained with other intensity range combinations. This suggests that stimulus range affects the perception of loudness in addition to whatever effects it may have on numerical judgments of loudness.     

Top-down gain control in the auditory system: evidence from identification and discrimination experiments

The influence of intensity range in auditory identification and intensity discrimination experiments is well documented and is usually attributed to nonsensory factors. Recent studies, however, have suggested that the stimulus range effect might be sensory in origin. To test this notion, in one set of experiments, we had listeners identify the individual tones in a set. One baseline condition consisted of identifying four 1-kHz, low-intensity tones; the other consisted of identifying four 1-kHz, high-intensity tones. In the experimental conditions, these baseline tone sets were augmented by adding a fifth tone at either 1 or 5 kHz. Added 5-kHz tones had little effect on identification accuracy for the four baseline tones. When an added 1-kHz tone differed substantially in intensity from the four baseline tones, it adversely affected performance, with the addition of a high-intensity tone to a set of low-intensity tones having a more deleterious effect than the addition of a low-intensity tone to a set of high-intensity tones. These and further results, obtained in an exploration of this asymmetrical range effect in a third identification experiment and in two intensity-discrimination experiments, were consistent with the notion of a nonlinear amplifier under top-down control whose functions include protection against sensory overload from loud sounds. The identification data were well described by a signal-detection model using equal-variance Laplace distributions instead of the usual Gaussian distributions.     

Name that percussive tune: Associative memory and amplitude envelope†

A series of experiments demonstrated novel effects of amplitude envelope on associative memory, with tones exhibiting naturally decaying amplitude envelopes (e.g., those made by two wine glasses clinking) better associated with target objects than amplitude-invariant tones. In Experiment 1 participants learned associations between household objects and 4-note tone sequences constructed of spectrally matched pure tones with either “flat” or “percussive” amplitude envelopes. Those hearing percussive tones correctly recalled significantly more sequence–object associations. Experiment 2 demonstrated that participants hearing percussive tones learned the associations more quickly. Experiment 3 used “reverse percussive” tones (percussive tones played backwards) to test whether differences in overall energy might account for this effect, finding they did not lead to the same level of performance as percussive tones. Experiment 4 varied the envelope at encoding and retrieval to determine which stage of the task was most affected by the envelope manipulation. Participants hearing percussive tones at both encoding and retrieval performed significantly better than the other three groups (i.e., flat at encoding/percussive at retrieval, etc.). We conclude that amplitude envelope plays an important role in learning and memory, a finding with relevance to psychological research on audition and associative memory, as well as practical relevance for improving human–computer interface design.     

Individual loudness functions determined from direct comparisons of loudness intervals

Five subjects were required in each trial to directly compare two pairs of tones and indicate which pair of tones had the greater loudness difference. Ten 1,200-Hz tones differing only in intensity were employed. Subjects made binary comparisons among the 45 tone pairs that can be formed from these 10 tones. The loudness difference comparisons of each subject were found to satisfy two properties (transitivity and monotonicity) that are required for an interval scale representation of loudness. Therefore, individual loudness scales were constructed using a nonmetric scaling technique designed for comparisons of sensory intervals. These loudness scales differed significantly from subject to subject. Since a nonnumerical scaling procedure was employed, these individual differences could not be attributed to biases in the way in which observers use numbers or numerical concepts to describe the loudness of tones. Hence, they suggest strong individual differences in the coding of sound intensity.     

Time errors and differential sensation weighting

Sixteen pairs of successive tones, with different amplitude combinations, were presented with 16 combinations of tone duration and interstimulus interval. A separate group of 12 subjects was assigned to each presentation condition and made comparative loudness judgments for each of the pairs. Perceived within-pair loudness differences were scaled by a Thurstonian method using the subjective width of the "equal" category as the unit. The scale differences were well described by weighted linear combinations of the sensation magnitudes of the tones in the pairs. The time error can be regarded as an effect of this differential weighting. For the longer interstimulus intervals, the weight of the second tone was the greater, causing the usual inverse relation between time error and stimulus intensity level. For the shorter interstimulus intervals, these effects were reversed. An analysis of the pattern of weights led to the development of two models, one of which is a generalization of Michels and Helson's time error model. The weights could be interpreted as reflecting the differential efficiency of the loudness information from the two compared stimuli.     

Accents in equitone sequences

Equitone sequences are defined as sequences of tones that are identical in all respects: frequency, spectral composition, intensity, and duration. The only parameter varied in these sequences is the time-interval between tones. In such sequences, clear accents are perceived. This paper describes accent perception in equitone sequences containing two alternating intervals; such sequences are perceived as consisting of repeating groups of two tones. An accent is heard on the first tone of a group if the difference between the intervals is about 5% to 10%. If the difference is made bigger, the accent is heard on the last tone of the group; this latter accent is considerably stronger than the accent previously heard on the first tone. In a number of experiments, the conditions under which the two types of accents occur were investigated. From these experiments, it was tentatively concluded that the accent on the last tone is heard because that tone, since it is followed by a longer interval, can be processed more completely. This “intervalproduced” accent indeed occurs only if the between-group interval is considerably longer than the within-group interval and if the latter does not exceed a duration of about 250 msec. The effect is slightly dependent on tone duration. The interval-produced accent can be balanced if the nonaccented tone is increased by about 4 dB in intensity. This shows that the effect is quite robust. The specific type of accentuation reported here might explain some rhythmical phenomena, examples of which are given.     

Generalization of cross-modal stimulus equivalence classes: operant processes as components in human category formation.

This study employed a stimulus-class rating procedure to explore whether stimulus equivalence and stimulus generalization can combine to promote the formation of open-ended categories incorporating cross-modal stimuli. A pretest of simple auditory discrimination indicated that subjects (college students) could discriminate among a range of tones used in the main study. Before beginning the main study, 10 subjects learned to use a rating procedure for categorizing sets of stimuli as class consistent or class inconsistent. After completing conditional discrimination training with new stimuli (shapes and tones), the subjects demonstrated the formation of cross-modal equivalence classes. Subsequently, the class-inclusion rating procedure was reinstituted, this time with cross-modal sets of stimuli drawn from the equivalence classes. On some occasions, the tones of the equivalence classes were replaced by novel tones. The probability that these novel sets would be rated as class consistent was generally a function of the auditory distance between the novel tone and the tone that was explicitly included in the equivalence class. These data extend prior work on generalization of equivalence classes, and support the role of operant processes in human category formation.     

Perceptual effects of preceding nonspeech rate on temporal properties of speech categories

The rate of context speech can influence phonetic perception. This study investigated the bounds of rate dependence by observing the influence of nonspeech precursor rate on speech categorization. Three experiments tested the effects of pure-tone precursor presentation rate on the perception of a [ba]-[wa] series defined by duration-varying formant transitions that shared critical temporal and spectral characteristics with the tones. Results showed small but consistent shifts in the stop-continuant boundary distinguishing [ba] and [wa] syllables as a function of the rate of precursor tones, across various manipulations in the amplitude of the tones. The effect of the tone precursors extended to the entire graded structure of the [w] category, as estimated by category goodness judgments. These results suggest a role for durational contrast in rate-dependent speech categorization.     

Temporal preparation decreases perceptual latency: Evidence from a clock paradigm

A clock paradigm was employed to assess whether temporal preparation decreases the time to detect the onset of a stimulus—that is, perceptual latency. In four experiments participants watched a revolving clock hand while listening to soft or loud target tones under high or low temporal preparation. At the end of each trial, participants reported the clock hand position at the onset of the target tone. The deviation of the reported clock hand position from the actual position indexed perceptual latency. As expected, perceptual latency decreased with target tone intensity. Most importantly, however, greater temporal preparation decreased perceptual latency in all four experiments, especially for soft tones, which supports rather directly the idea that temporal preparation diminishes the duration of perceptual processing.     

Contrast effects do not underlie effects of preceding liquids on stop-consonant identification by humans

These experiments explored the claim by A. Lotto and K. Kluender (1998) that frequency contrast explains listeners' compensations for coarticulation in the case of liquid consonants coarticulating with following stops. Evidence of frequency contrast in experiments that tested for it directly was not found, but Lotto and Kluender's finding that high- and low-frequency precursor tones can produce contrastive effects on stop-consonant judgments were replicated. The effect depends on the amplitude relation of the tones to the third formant (F3) of the stops. This implies that the tones mask F3 information in the stop consonants. It is unknown whether liquids and following stops in natural speech are in an appropriate intensity relation for masking of the stop. A final experiment, exploiting the McGurk effect, showed compensation for coarticulation by listeners when neither frequency contrast nor masking can be the source of the compensations.     

Intensity discrimination and loudness for tones in broadband noise

In two previous papers (Parker & Schneider, 1980; Schneider & Parker, 1987), we developed a model, based on Fechner's assumption, which successfully predicted the relationship between loudness and intensity discrimination for tones presented in quiet and in notched noise. In the present paper, pure-tone intensity-increment thresholds and loudness matches were determined for several levels of a standard tone in the presence of a broadband masker whose spectrum level was set to 35 dB below that of the standard tone. The model was unable to relate loudness to intensity discrimination under these conditions. Thus, the spectral composition of the masker affects the relationship between loudness and intensity discrimination in ways that cannot be accounted for by the model.     

Loudness and loudness discrimination

A model is developed which holds that pure-tone intensity discrimination and suprathreshold loudness judgments are based on the same sensory representation. In this model, loudness is a power function of sound intensity. When two tones are presented sequentially, each gives rise to a loudness value along the sensory continuum. In intensity-discrimination experiments, threshold is reached when the loudness difference between the tones exceeds a criterial value. For suprathreshold presentations of tone pairs, judgments of loudness differences are based on the loudness difference between the two tones. The model is shown to accord well with data from both classes of experiments.     

汉语语音范畴性知觉及其发展

范畴性是言语知觉的一个显著特征,长期以来一直受到研究者的普遍关注。但汉语不同语音特征的范畴性知觉特点及其发展模式迄今为止还很少有研究涉及。本研究利用语音合成的方法分别改变辅音的送气/不送气特征和声调的基频曲线,生成语音刺激连续体,采用范畴性知觉的经典研究范式探讨了汉语正常成人被试嗓音启动时间（Voice onset time,VOT）和声调范畴性知觉的特点及不同年龄儿童的发展模式。研究结果表明：（1）成人被试对VOT和声调的知觉是范畴性的;（2）对于汉语声调特征,6岁儿童已经具有类似成人的范畴性知觉能力;而在VOT这一维度上,范畴知觉能力随着年龄发展不断精细化,但7岁儿童也尚未达到成年人的敏感程度,说明汉语VOT和声调这两个不同的语音特征经历了不同的发展模式。